A motion of no confidence in Labour leader Jeremy Corbyn has been passed by the party's MPs.
The 172-40 vote, which is not binding, follows resignations from the shadow cabinet and calls on Mr Corbyn to quit.
Mr Corbyn said the ballot had "no constitutional legitimacy" and said he would not "betray" the members who voted for him by resigning.
The leader's allies have told his critics to trigger a formal leadership contest if they want to challenge him.
There were four abstentions in Labour's secret ballot.
Labour's leader in Scotland, Kezia Dugdale, suggested Mr Corbyn's position was untenable, telling the BBC: "If I had lost the support of 80% of my MSPs I could not do my job."
Labour MP and Corbyn critic Wes Streeting said the vote was "unprecedented", adding: "I think Jeremy just has to accept now that his leadership is untenable."
But following the result the leader issued a statement saying the government was "in disarray" following the vote to leave the EU, adding: "Labour has the responsibility to give a lead where the government will not."
He added: "I was democratically elected leader of our party for a new kind of politics by 60% of Labour members and supporters, and I will not betray them by resigning. Today's vote by MPs has no constitutional legitimacy.
"We are a democratic party, with a clear constitution. Our people need Labour Party members, trade unionists and MPs to unite behind my leadership at a critical time for our country."

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